Steve Borthwick warned England must be prepared "go through some pain" after they were beaten by Scotland in his first game in charge at Twickenham on Saturday.

There was no dream start for Borthwick after he replaced the sacked Eddie Jones as Red Rose head coach in December, as Scotland retained the Calcutta Cup with a 29-23 win.

Duhan van der Merwe starred in an entertaining Six Nations clash, scoring a brilliant solo try in the first half and adding a second five minutes from time to give Gregor Townsend's side a bonus-point victory.

That is three consecutive wins for Scotland over England, who led by eight points early in the second half but were unable to see it out.

Borthwick said: "This is part of the growth of the team.

"We got ourselves in a position when we should have won the Test match in the second half. At 20-12 up, we shouldn't be letting that game get away from us. You have to go through some pain.

"Really good teams would go on and win that.

"We were playing against a Scotland team who have controlled this fixture in recent years. We need to make sure we limit chances and can shut them down."

Borthwick says it is not all doom and gloom after a losing start to his reign.

He added: "We saw some growth, particularly in the attacking side of the game. The game was quicker, but clearly we're disappointed in the result.

"The team in the autumn, if they conceded a couple of scores early didn't come back from that. These guys did.

"Unfortunately we let the opposition back in. We want to be a really successful team that wins Test matches. When you get to that level, you don't give the opposition the opportunities we did.

"I am disappointed for the supporters, who I thought were incredible for us today. We'll make sure we come back here in eight days' time and are a better team."